Datta et al. Appendix S2 Appendix S2. Parasuchus hislopi Lydekker (1885). A, neotype, ISIR 42, actual photograph of a nearly complete skull in right lateral view. B–C, boxed area I of A at higher magnification, showing right lateral surface of the jugal between the antorbital and the infratemporal fenestra; B, actual photograph; C, line drawing. D, neotype, ISIR 42, actual photograph of a nearly complete skull in left lateral view. E–F, boxed area II of D at higher magnification, showing left lateral surface of the jugal between the antorbital and the infratemporal fenestra; E, actual photograph; F, line drawing. Abbreviations: afn, antorbital fenestra; afo, antorbital fossa; asc, ascending maxillary process; dsc, descending maxillary process; ec, ectopterygoid; en, external nare; f, frontal; itf, infratemporal fenestra; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pmx, premaxilla; prf, prefrontal; q, quadrate; qj, quadratojugal. Scale bars represent 30 mm. Datta et al._Appendix S3 Appendix S3. Cranial characters used in the phylogenetic analysis. A brief explanation follows the new characters used, whereas for the characters already used in previous literature, its citation follows the last character state of each. The citations are abbreviated as follows: H, Hungerbühler (2002); S, Stocker (2010); K, Kammerer et al. (2016); J, Jones & Butler (2018), whereas the numerals indicate the character number of each citation. Datta et al._Appendix S4 Appendix S4. Character matrix used for phylogenetic analysis. Datta et al_Appendix S5 Appendix S5. TNT file of Volcanosuchus statisticae. Datta et al._Appendix S1 Appendix S1. Volcanosuchus statisticae gen et sp. nov. Measured parameters (in mm).

Detailed description and phylogenetic assessment of a phytosaur skull collected from the Tiki Formation of the Rewa Gondwana Basin of India and earlier diagnosed as Parasuchus hislopi, shows that it pertains to a new genus and species, Volcanosuchus statisticae. The new taxon is characterized by marginal overlapping of the nostrils by the antorbital fenestrae, external nares situated on a bulbous and raised dome, the lateral surface of the jugal ornamented by a prominent ridge defined by multiple tubercles and radiating thread-like structures, and distinct ornamentation patterns on the rostrum and skull table. Phylogenetic analysis nests Volcanosuchus within Mystriosuchinae where it forms a sister taxon to (Rutiodon + Leptosuchomorpha) and marks the transition between the basal Parasuchidae and more derived Mystriosuchinae phytosaurs. Evolution of the phytosaur skulls resulted in changes from non-overlapping nostril and antorbital fenestra to an overlapping state, anteroposterior elongation of the exoccipital and supraoccipital shelf, appearance of a median ridge on the basioccipital, and reduction of the supratemporal fenestra. Considerable faunal overlap of the Tiki Formation is evident with the lower Maleri Formation, which is late Carnian based on Hyperodapedon, Parasuchus, and Exaeretodon. The Tiki Formation correlates with the Ischigualasto Formation of Argentina, the upper part of the Santa Maria Formation, and the overlying lower Caturrita Formation of Brazil, the Isalo II Beds of Madagascar, Lossiemouth Sandstone of Scotland, and the lower Tecovas Formation of the Chinle Group of North America, and ranges from late Carnian to early/middle Norian.