Four new species of Passiflora from Ecuador are described, viz. P. montana, P. palenquensis, P. deltoifolia, and P. pergrandis. A revision of Ecuadorean Passiflora subg. Passiflora (subg. Granadilla sensu Killip) for the Flora of Ecuador (Holm-Nielsen et al., in press) has revealed four new species: P. montana, P. palenquensis, P. deltoifolia, and P. pergrandis, belonging to "series" Lobatae, Tiliaefoliae, Menispermifoliae, and Laurifoliae, respectively. The total number of Ecuadorean subg. Passiflora species is 19. Killip (1938) divided the subgenus into 15 series; although these are not validly published, we are following Killip for convenience. The "series" occurring in Ecuador are Digitatae, Incarnatae, Kermesianae, Simplicifoliae, Quadrangulares, and those mentioned above. Passiflora palenquensis Holm-Nielsen & Lawesson, sp. nov. TYPE: Ecuador. Los Rios: Quevedo-Sto. Domingo road, km 56, Rio Palenque Science Center, 150-200 m, 6 Oct. 1979, C. H. Dodson, A. Gentry & G. Shupp 8854 (holotype, MO). Figure 1. Liana ubique glabra. Caule striato et tereto. Stipulis oblongo-lanceolatis, acutis, 1 x 0.4 cm, interdum decideosis. Petiolis angulo-alatis, 6.5-9 cm longis cum 3-4 nigris, sessilibus urceolatiaque glandibus, par apice extremo, aliae glandes infra 2-3 cm; laminibus basifixis, late ovatis, ovatis, 14-19 x 11-20 cm, profunda cordatis, manifeste pinnatinervibus, nitentibus coriaceis, margine integra. Inflorescentia saltem 2 floribus; pedicellis teretis striatisque, 3-4 cm longis; bracteae circa 5 mm infra florem portatae, verticillatae, connatae dimidium, oblongo-ovatae, mucronatae, 5 x 2 cm. Floribus 5-6 cm latis; hypanthium infundibulare, longo 3 cm, ad basin 1 cm lato, ad apicem 2 cm, lavandulo; lobi calycis oblongi, acuti, 2.5 x 1.3 cm, desinentes in minutam mucronem; petalis oblongis, obtusis, 3 x 0.6-1 cm. Corona simplice, filamentis composita in 2 seriebus ordinatis quae ad apicem extremam tubi calycis locatae sunt; series externa minute liguliformis, apex filiformis, 4 mm longa; serie interna crassisima, 4-angulata, 2 cm longa, apex filiformis, fasciata purpurea et alba; tubus calycis glaber infra coronam. Operculum positum circa 1 cm supra pavimentum tubi calycis, membranaceum, in superiore parte filiforme, 0.5 cm longum. Inferioris partis 0.5 cm gynophori limini circumcinctae. Ovario glabro. Liana, glabrous throughout. Stem terete, striate. Stipules oblong-lanceolate, acute, 1 x 0.4 cm, sometimes deciduous. Petioles angular-winged, 6.5-9 cm long, with 3-4 black, sessile urceolate glands, 1 mm long, one pair at extreme apex, other glands 2-3 cm below; blades basifixed, broad-ovate to ovate, 14-19 x 1 1-20 cm, deeply cordate, prominently pinnate-veined, lustrous, coriaceous, the margin entire. Inflorescence sessile, with at least 2 flowers collateral with the tendril in the axil of the leaf, 5-6 cm diam.; pedicels terete, striate, 3-4 cm long; bracts verticillate, united halfway, inserted about 5 mm below flower, oblong-ovate, mucronate, 5 x 2 cm. Hypanthium tube funnel-shaped, 3 cm long, 1 cm wide at base, 2 cm at apex, outside lavender; calyx lobes oblong, acute, 2.5 x 1.3 cm, ending in a minute mucro; petals oblong, obtuse, 3 x 0.6-1 cm. Corona composed of 2 series situated at the edge of the hypanthium; outer series of short ligulate filaments, 4 mm long, with filiform apices; inner series of very stout, 4-angled, 2 cm long filaments with filiform apices, the inner filaments with shifting purple and white, 4 mm broad stripes; hypanthium smooth below corona. Operculum situated 1 cm above hypanthium floor, membranaceous, 0.5 cm long, split into filiform segments. Trochlea present at the androgynophore opposite the apex of the operculum. Limen surrounding lower 5 mm of androgynophore. Gynophore 1 cm longer than the androgynophore present. Ovary glabrous. Fruit I We thank R. R. Haynes and H. Balslev for reading the manuscript, K. Tind for making the drawings, and N.-H. Andreasen, who supplied the Latin diagnoses. Fieldwork was supported by the Danish Natural Science Research Council, grant numbers 11-3038, 11-3689, and 81-4024. This is contribution No. 53 from the AAU Ecuador Project. 2 Botanical Institute, University of Aarhus, 68 Nordlandsvej, DK-8240 Risskov, Denmark. 3 Charles Darwin Research Station Galapagos, Casilla 5839, Guayaquil, Ecuador. ANN. MissouRI BOT. GARD. 74: 497-504. 1987. This content downloaded from 207.46.13.129 on Sun, 26 Jun 2016 05:32:49 UTC All use subject to http://about.jstor.org/terms 498 ANNALS OF THE MISSOURI BOTANICAL GARDEN [VOL. 74